Morphology of the Roots – PST05210 Basic Pharmacognosy

NTA Level 5 • Semester 2 • PST05210

Morphology of the Roots

Basic Pharmacognosy • Source Session/Topic 5
Full source-text version: all educational wording from the extracted learning source is retained; only presenter/tutor metadata and web-layout noise are removed, while formatting is improved for readability.

Session 5: Morphology of the Roots

Total Session Time: 120 minutes

Prerequisites

• None

Learning Tasks

By the end of this session students are expected to be able to:

• Explain Roots
• Classify Roots
• Differentiate Monocotyledon from Dicotyledon
• Functions of the Roots

Resources Needed:

• Flip charts, marker pens, and masking tape
• Black/white board and chalk/whiteboard markers

SESSION OVERVIEW

|Step |Time |Activity/ |Content |

| | |Method | |

|1 |05 minutes |Presentation |Introduction, Learning Tasks |

|2 |25 minutes |Presentation |Characteristics of Plant |

| | | |Roots |

|3 |30 minutes |Presentation |Anatomy of Monocotyledonous |

| | | |Roots |

|4 |30 minutes |Presentation |Anatomy of Dicotyledonous |

| | | |Roots |

|5 |20 minutes |Presentation |Difference between Monocot |

| | |Small group |and Dicot Roots |

| | |Discussion | |

|6 |05 minutes |Presentation |Key Points |

|7 |05 minutes |Presentation |Evaluation |

SESSION CONTENTS

STEP 1: Presentation of Session Title and Learning Tasks (5 minutes)

READ or ASK students to read the learning objectives and clarify

ASK students if they have any questions before continuing

STEP 2: Characteristics of Plant Roots (25 minutes)

• The root constitutes the underground part of the plant axis.
• It is an organ of the plant that typically lies below the surface of

the soil

• Externally the root has root hairs which aid in absorption of water

minerals

• Roots can be classified in three main categories:

o Tap roots

▪ One large vertical root
▪ Also produces many small lateral or branch roots
▪ These are common in dicotyledonous plants

o Fibrous roots

▪ Mat of thin roots that spread out
▪ Made up of many thread-like members of more or less equal length
▪ These are common in monocotyledonous plants

o Adventitious roots

▪ Form on any plant part other than the roots
▪ They commonly occur in grasses
• General Internal structure of the root;

[pic]

• Functions of the Roots

o The following are the functions of the roots

▪ Roots support (anchorage) the plant in the soil (or other media in

which the plant is growing)

▪ Absorption of water and other nutrients from the soil
▪ Storage of food
▪ Transport minerals and water to other parts of the plant

STEP 3: Anatomy of the Monocotyledonous Roots (30 minutes)

• Internal structure of monocot roots shows the following tissue systems

o Epiblema or rhizodermis

o Cortex and

o Stele

Fig 5.1: Internal Structure of the Monocotyledonous Root

[pic]

• The Rhizodermis or Epiblema

o Outermost layer of the root

o Single row of parenchyma cells, no intercellular space

o Stomata and cuticle are absent

o Root hairs are always unicellular

o The key function of rhizodermis is protection of the inner

tissues of the root

• The Cortex

o The cortex is made of only one type of tissue (parenchyma)

o Has many layers of parenchyma cells with lot of intercellular spaces

o Cortical cells are generally oval or rounded in shape; chloroplasts are

absent

o The cells are living and possess leucoplasts

o The inner most layer of the cortex is endodermis and is composed of

single layer of barrel shaped parenchymatous cells

o The function of cortical cells is storage

o Endodermis (innermost layer of cortex) forms a complete ring around the

stele (tissues inside endodermis)

o There is a band-like structure made of (thickened with) suberin present

in the radial and transverse walls of the endodermal cells

o The band is called Casparian strips

o The endodermal cells opposite to the protoxylem elements, are thin-

walled without Casparian strips

o These cells are called passage cells and their function is to transport

water and dissolved salts from the cortex to the xylem

o Water cannot pass through other endodermal cells due to Casparian

strips (blocking effect)

o The main function of Casparian strips in the endodermal cells is to

prevent the re-entry of water into the cortex once water enters the

xylem tissue

• Vascular System (Stele)

o Vascular tissues are in radial arrangement with many protoxylem

groups

o This arrangement of xylem is called polyarch and the xylem is in

exarch condition

o The tissue present between the xylem and the phloem, is called

conjunctive tissue

• The Pith

o The central portion

o It consists of thin walled parenchyma cells with intercellular

spaces

o These cells are filled with abundant starch grains

STEP 4: Anatomy of the Dicotyledonous Roots (30 minutes)

• The Dicotyledonous Root

o Structure of a Dicotyledonous Root

[pic]

• Rhizodermis or Epiblema

o Outermost layer aka rhizodermis (epidermis in stems) made of a single

layer of parenchyma cells which are arranged compactly without

intercellular spaces, with no stomata or cuticle

o Root hair is always single celled

o Protects inner tissues of the root

• The Cortex

o Consists of only parenchyma cells loosely arranged with intercellular

spaces to make gaseous exchange easier

o These cells may store food reserves

o The cells are oval or rounded in shape

o Sometimes are polygonal due to mutual pressure

o Cortical cells are devoid of chloroplasts but starch grains are stored

in them

o The cortical cells also possess leucoplasts

o Inner most layer of the cortex is endodermis

o Endodermis is made up of single layer of barrel shaped parenchymatous

cells

o Stele is completely surrounded by the endodermis

o The radial and the inner tangential walls of endodermal cells are

thickened with suberin

o Casparian strips

o Casparian strips are absent in the endodermal cells located opposite to

the protoxylem elements

o These thin-walled cells without Casparian strips are called passage

cells through which water and mineral salts are conducted from the

cortex to the xylem elements

o Water cannot pass through other endodermal cells due to the presence of

Casparian thickenings

• The Stele

o Include all the tissues present inside endodermis

o The stele includes pericycle and vascular tissues

▪ The Pericycle
• Pericycle is generally a single layer of parenchymatous cells found

inner to the endodermis

• It is the outermost layer of the stele
• Lateral roots originate from the pericycle
• Thus, the lateral roots are endogenous in origin
▪ The Vascular Tissues
• Vascular tissues are in radial arrangement
• Xylem and phloem are separated by a conjunctive tissue
• If conjunctive tissue is composed of parenchymatous tissue then,

the xylem is in exarch condition

• If number of protoxylem points is four, then, the xylem is called

tetrarch

• Each phloem patch consists of sieve tubes, companion cells and

phloem parenchyma

• Metaxylem vessels are generally polygonal in shape but in monocot

roots they are circular

STEP 5: Difference between Monocot and Dicot Roots (20 minutes)

|Activity: Small Group Discussion (15 minutes) |

| |

|DIVIDE students into small manageable groups |

| |

|ASK students to discuss on the following question |

|What are the differences between monocot root and dicot root? |

| |

|ALLOW students to discuss for 15 minutes |

| |

|ALLOW few groups to present and the rest to add points not |

|mentioned |

| |

|CLARIFY and SUMMARIZE by using the contents below |

[pic]

STEP 6: Key Points (10 minutes)

• The root is an organ of the plant that typically lies below the surface

of the soil

• Three basic types of roots include; tap roots, fibrous roots and

adventitious roots

• Anatomy of the root consists of a series of concentric rings of different

tissues namely the epidermis, the cortex, the endodermis, the pericyle

and the vascular system

STEP 7: Evaluation (5 minutes)

• What is a root
• What are the layers composing the internal structure of the dicot root?

References

Trease, G. E., Evans, W. C., & Evans, D. (2009). Trease and Evans

pharmacognosy. London: Saunders.

Joanne Barnes et al (2002), Herbal medicines 3rd Edition: Pharmaceutical

Press

Wallis, T. E. (2005). Textbook of pharmacognosy. New Delhi: CBS.

Robbers, J. E., Speedie, M. K., Tyler, V. E., & Tyler, V. E. (1996).

Pharmacognosy and pharmacobiotechnology. Baltimore: Williams &

Wilkins.

Heinrich, M., Kinghorn, A. D., Maizels, D., Gibbons, S., & Phillipson, J.

D. (2012). Fundamentals of pharmacognosy and phytotherapy. Edinburgh:

Churchill Livingstone/Elsevier.

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